Physical Sciences and Mathematics Commons^™

Cytoarchitectural Characteristics Associated With Cognitive Flexibility In Raccoons, Joanna Jacob, Molly Kent, Sarah Benson-Amram, Suzana Herculano-Houzel, Mary Ann Raghanti, Emily Ploppert, Jack Drake, Bilal Hindi, Nick R. Natale, Sarah Daniels, Rachel Fanelli, Anderson Miller, Tim Landis, Amy Gilbert, Shylo Johnson, Annie Lai, Molly Hyer, Amanda Rzucidlo, Chris Anchor, Stan Gehrt, Kelly Lambert

United States Department of Agriculture Wildlife Services: Staff Publications

With rates of psychiatric illnesses such as depression continuing to rise, additional preclinical models are needed to facilitate translational neuroscience research. In the current study, the raccoon (Procyon lotor) was investigated due to its similarities with primate brains, including comparable proportional neuronal densities, cortical magnification of the forepaw area, and cortical gyrification. Specifically, we report on the cytoarchitectural characteristics of raccoons profiled as high, intermediate, or low solvers in a multiaccess problem-solving task. Isotropic fractionation indicated that high-solvers had significantly more cells in the hippocampus (HC) than the other solving groups; further, a nonsignificant trend suggested that this …

Movement, Space-Use And Resource Preferences Of European Golden Jackals In Human-Dominated Landscapes: Insights From A Telemetry Study, Skye Fenton, Paul R. Moorcroft, Duško Ćirović, József Lanszki, Miklós Heltai, Francesca Cagnacci, Stewart Breck, Neda Bogdanović, Ilija Pantelić, Kornél Ács, Nathan Ranc

United States Department of Agriculture Wildlife Services: Staff Publications

During the last half-century, the distribution of golden jackals (Canis aureus) has rapidly increased throughout Europe. Today, golden jackals are thriving in human-dominated landscapes across Southeastern and Central Europe. Most studies on golden jackals have focused on large-scale distribution patterns; to date, little is known about the species’ fine-scale spatial ecology. In this study, we analyzed the movement behavior, space-use and resource selection of six golden jackals fitted with GPS-GSM collars in two study areas in Hungary and Serbia. Two of the jackals were a breeding pair. We found that home range size averaged 11.2 km² (90% …

Threading The Needle: How Humans Influence Predator–Prey Spatiotemporal Interactions In A Multiple-Predator System, Asia Murphy, Duane R. Diefenbach, Mark Ternent, Matt Lovallo, David Miller

United States Department of Agriculture Wildlife Services: Staff Publications

Perceived predation risk and the resulting antipredator behaviour varies across space, time and predator identity. Communities with multiple predators that interact and differ in their use of space, time of activity and hunting mode create a complex landscape for prey to avoid predation. Anthropogenic presence and disturbance have the potential to shift interactions among predators and prey and the where and when encounters occur. We examined how white-tailed deer Odocoileus virginianus fawn spatiotemporal antipredator behaviour differed along an anthropogenic disturbance gradient that had black bears Ursus americanus, coyotes Canis latrans, bobcats Lynx rufus and humans present. We quantified (a) spatial …

Virginia Seafood Sustainability, Samantha E. Askin, Robert A. Fisher

Reports

Virginia’s commercial fisheries operate sustainability under a suite of management tools based upon information received from marine scientists and fishery managers who regularly conduct biological sampling of fish while tracking commercial landings and other gathering of required information. Analyses of fishing effort and overall stock conditions, as well as formulas designed to calculate threshold limits for maintaining sustainable stocks are regularly performed. Restrictions on seasons, size, days at sea, and gear are imposed as needed to achieve management supporting long-term biological sustainability.

The Pandemic, Climate Change And Farm Subsidies, Allen H. Olson, Edward J. Peterson

Journal of Food Law & Policy

Many people believe that once the COVID-19 pandemic has passed, life will return to the way it was. This belief is both unrealistic and dangerous. It is unrealistic because the virus will be around for years if not indefinitely. The timeframe for the worst of the pandemic will depend on our ability to administer effective vaccines worldwide and the public’s willingness to accept continued social distancing in the meantime. The damage done to public health, the economy and individuals is already substantial and will get worse. Recovery will be slow and incomplete. The belief that life will return to the …

Socio-Economic And Governance Conditions Corresponding To Change In Animal Agriculture: South Dakota Case Study, Jacqueline S. Welles, Noelle Cielito T. Soriano, Freda Elikem Dorbu, G M. Pereira, Laura M. Rubeck, Erica L. Timmermans, Benjamin Ndayambaje, Alison V. Deviney, John J. Classen, Jacek A. Koziel, Erin L. Cortus

School of Natural Resources: Faculty Publications

Understanding sustainable livestock production requires consideration of both qualitative and quantitative factors in a temporal and/or spatial frame. This study adapted Qualitative Comparative Analysis (QCA) to relate conditions of social, economic, and governance factors to changes in livestock inventory across several counties and over time. This paper presents an approach that (1) identified factors with the potential to relate to a change in livestock inventory and (2) analyzed commonalities within these factors related to changes spatially and temporally. This paper illustrates the approach and results when applied to five counties in eastern South Dakota. The specific response variables were periods …

Based On Atmospheric Physics And Ecological Principle To Assess The Accuracies Of Field Co2 /H2o Measurements From Infrared Gas Analyzers In Closed-Path Eddy-Covariance Systems, Xinhua Zhou, Tian Gao, Yunchao Pang, Hayden Mahan, Xiufen Li, Ning Zheng, Andrew E. Suyker, Tala Awada, Jiaojun Zhu

School of Natural Resources: Faculty Publications

Field CO₂ /H₂O measurements from infrared gas analyzers in closed-path eddy-covariance systems have wide applications in earth sciences. Knowledge about exactness of these measurements is required to assess measurement applicability. Although the analyzers are specified with uncertainty components (zero drift, gain drift, cross-sensitivities, and precision), exactness for individual measurements is unavailable due to an absence of methodology to comprehend the components as an overall uncertainty. Adopting an advanced definition of accuracy as a range of all measurement uncertainty sources, the specified components are composited into a model formulated for studying analyzers’ CO₂ /H₂O accuracy …

The Relative Importance Of Wind-Driven And Chimney Effect Cave Ventilation: Observations In Postojna Cave (Slovenia), Lovel Kukuljan, Franci Gabrovsek, Matthew Covington

International Journal of Speleology

Density-driven chimney effect airflow is the most common form of cave ventilation, allowing gas exchange between the outside and the karst subsurface. However, cave ventilation can also be driven by other mechanisms, such as barometric changes or pressure differences induced by the outside winds. We discuss the mechanism and dynamics of wind-driven ventilation using observations in Postojna Cave, Slovenia. We show how seasonal airflow patterns driven by the chimney effect are substantially modified by outside winds. Wind flow over irregular topography forms near-surface air pressure variations and thus pressure differences between cave entrances at different locations. These pressure differences depend …

A Systems Approach To Understanding Piping Plover Production On Off-Channel Sandpits Along The Central Platte River, Taylor Morgan Cassidy

Biology Theses, Dissertations, and Student Creative Activity

Anthropogenic changes in land use, exploitation of natural resources, climate change, and habitat loss create difficulties managing and conserving species of concern. Managers are often forced to make decisions with incomplete information and much uncertainty. Unsuccessful attempts to manage for and conserve species in the past, may have resulted from failures to recognize the complexity of systems, resulting in actions that may have had unintended consequences. Thus, applying Systems Thinking and System Dynamics is appropriate for understanding what factors influence species success and evaluating what management actions may be implemented to help achieve conservation targets while minimizing negative impacts to …

A Proposed Framework For The Development And Qualitative Evaluation Of West Nile Virus Models And Their Application To Local Public Health Decision-Making, Alexander C. Keyel, Morgan E. Gorris, Ilia Rochlin, Johnny A. Uelmen, Luis F. Chaves, Gabriel L. Hamer, Imelda K. Moise, Marta Shocket, A Marm Kilpatrick, Nicholas B. Defelice, Justin K. Davis, Eliza Little, Patrick Irwin, Andrew J. Tyre, Kelly Helm Smith, Chris L. Fredregill, Oliver Elison Timm, Karen M. Holcomb, Michael C. Wimberly, Matthew J. Ward, Christopher M. Barker, Charlotte G. Rhodes, Rebecca L. Smith

School of Natural Resources: Faculty Publications

West Nile virus(WNV) is a globally distributed mosquito-borne virus of great public health concern. The number of WNV human cases and mosquito infection patterns vary in space and time. Many statistical models have been developed to understand and predict WNV geographic and temporal dynamics. However, these modeling efforts have been disjointed with little model comparison and inconsistent validation. In this paper, we describe a framework to unify and standardize WNV modeling efforts nationwide. WNV risk, detection, or warning models for this review were solicited from active research groups working in different regions of the United States. A total of 13 …

Underground Burning Of Jharia Coal Mine (India) And Associated Surface Deformation Using Insar Data, Jungrack Kim, Shih-Yuan Lin, Ramesh P. Singh, Chen-Wei Lan, Hye-Won Yun

Biology, Chemistry, and Environmental Sciences Faculty Articles and Research

The underground burning in the Jharia coal mine (JCM) in India is a highly devastating environmental hazard inducing various adverse consequences. In the present study, we carried out time series analyses based on Interferometric Synthetic Aperture Radar (InSAR) and land surface temperature (LST) to study the environmental risk. First, a permanent scatterer (PS) time series analysis using Sentinel-1 images over three years was performed to detect the spatio-temporal distribution of ground deformation. Comparison of ground thermal anomaly clearly delineated the subsidence spots associated with the oxygen supply to combustion areas. On the contrary, few deformations were mapped showing pronounced uplift …

Evaluation Of Regional Climate Models (Rcms) Using Precipitation And Temperature-Based Climatic Indices: A Case Study Of Florida, Usa, Yared Bayissa, Assefa Melesse, Mahadev Bhat, Tsegaye Tadesse, Andualem Shiferaw

School of Natural Resources: Faculty Publications

The overarching objective of this study was to evaluate the performance of nine precipitation-based and twelve temperature-based climatic indices derived from four regional climate models (CRCM5-UQUAM, CanRCM4, RCA4 and HIRHAM5) driven by three global circulation models (CanESM2, EC-EARTH and MPI-ESM-LR) and their ensemble mean for the reference period of 31 years (1975–2005). The absolute biases, pattern correlation, the reduction of variance (RV) and the Standardized Precipitation Evapotranspiration Index (SPEI at 3-, 6- and 12-month aggregate periods) techniques were used to evaluate the climate model simulations. The result, in general, shows each climate model has a skill in reproducing at least …

Plague Exposure In Mammalian Wildlife Across The Western United States, Sarah N. Bevins, Jeffrey C. Chandler, Nicole Barrett, Brandon S. Schmit, Gerald W. Wiscomb, Susan A. Shriner

United States Department of Agriculture Wildlife Services: Staff Publications

Plague is caused by a bacterial pathogen (Yersinia pestis) that can infect a wide range of mammal species, but its presence in wildlife is often underappreciated. Using a large-scale data set (n = 44,857) that details the extent of Y. pestis exposure in wildlife, we document exposure in 18 wildlife species, including coyotes (Canis latrans), bobcats (Lynx rufus), and black bears (Ursus americanus). Evidence of plague activity is widespread, with seropositive animals detected in every western state in the contiguous United States. Pathogen monitoring systems in wildlife that are both large scale …

Otoliths Of South-Western Australian Fish: A Photographic Catalogue, Chris Dowling, Kim Smith, Elaine Lek, Joshua Brown

Fisheries research reports

Due to the species-specific nature of otoliths and given they are often the only part of the fish preserved when fish die, otolith catalogues can be used in numerous applications, such as diet studies in fish eating animals, including pinnipeds, fish and sea birds; archaeological purposes such as reconstructing indigenous people’s diets from otoliths found in middens or evolutionary history of fish species by comparing fossilized otoliths. Given the unique mixture of subtropical and temperate fish, including many endemic species that occur off the southwest corner of WA having a catalogue for this area is extremely important for people working …

Winter Harbor Dredge Channel Data Report, Donna A. Milligan, C. Scott Hardaway Jr., Christine A. Wilcox, Cameron Green

Reports

Winter Harbor is a tidal creek that flows between a sandy barrier system and an eroding marsh shoreline. It is an important waterway that provides access to the Chesapeake Bay from the Winter Harbor watershed. In the past, Winter Harbor Inlet was the only hydraulic connection from the Winter Harbor watershed and Chesapeake Bay. The two open water areas of Winter Harbor were separated by marsh channels that hydraulically connected to the mouth at the present-day Winter Harbor Inlet. The federally-defined channel at Winter Harbor was authorized by Congress in 1950. It was authorized as a 12 feet (ft) deep, …

Improving Animal Monitoring Using Small Unmanned Aircraft Systems (Suas) And Deep Learning Networks, Meilun Zhou, Jared A. Elmore, Sathishkumar Samiappan, Kristine O. Evans, Morgan Pfeiffer, Bradley F. Blackwell, Raymond B. Iglay

United States Department of Agriculture Wildlife Services: Staff Publications

In recent years, small unmanned aircraft systems (sUAS) have been used widely to monitor animals because of their customizability, ease of operating, ability to access difficult to navigate places, and potential to minimize disturbance to animals. Automatic identification and classification of animals through images acquired using a sUAS may solve critical problems such as monitoring large areas with high vehicle traffic for animals to prevent collisions, such as animal-aircraft collisions on airports. In this research we demonstrate automated identification of four animal species using deep learning animal classification models trained on sUAS collected images. We used a sUAS mounted with …

Scaup Depredation On Arkansas Baitfish And Sportfish Aquaculture, Stephen A. Clements, Brian S. Dorr, J. Brian Davis, Luke A. Roy, Carole R. Engle, Katie C. Hanson-Dorr, Anita M. Kelly

United States Department of Agriculture Wildlife Services: Staff Publications

Lesser scaup (Aythya affinis) and greater scaup (A. marila), hereafter scaup, consume a variety of aquatic invertebrates, plants, and occasionally small fish. Scaup have foraged on commercial aquaculture farms in the southern United States for decades. However, the types, abundance, and rate of fish exploitation by scaup on baitfish and sportfish farms are not well documented. Thus, information is needed to understand how fish and other foods influence scaup use of aquatic resources, and any potential economic effects of depredation of fish. From November–March in winters 2016–2017 and 2017–2018, we conducted 1,458 pond surveys to estimate …

Data-Driven Management—A Dynamic Occupancy Approach To Enhanced Rabies Surveillance Prioritization, Amy J. Davis, Jordana D. Kirby, Richard B. Chipman, Kathleen M. Nelson, Amy T. Gilbert

United States Department of Agriculture Wildlife Services: Staff Publications

Rabies lyssavirus (RABV) is enzootic in raccoons across the eastern United States. Intensive management of RABV by oral rabies vaccination (ORV) has prevented its spread westward and shown evidence of local elimination in raccoon populations of the northeastern US. The USDA, Wildlife Services, National Rabies Management Program (NRMP) collaborates with other agencies to implement broad-scale ORV and conducts extensive monitoring to measure the effectiveness of the management. Enhanced Rabies Surveillance (ERS) was initiated during 2005 and updated in 2016 to direct surveillance efforts toward higher-value specimens by assigning points to different methods of encountering specimens for collection (strange-acting, roadkill, surveillance-trapped, …

Spatial Variation In Direct And Indirect Contact Rates At The Wildlife-Livestock Interface For Informing Disease Management, Anni Yang, Raoul K. Boughton, Ryan S. Miller, Bethany Wight, Wesley M. Anderson, James C. Beasley, Kurt C. Vercauteren, Kim M. Pepin, George Wittemyer

United States Department of Agriculture Wildlife Services: Staff Publications

Little is known about disease transmission relevant contact rates at the wildlife-livestock interface and the factors shaping them. Indirect contact via shared resources is thought to be important but remains unquantified in most systems, making it challenging to evaluate the impact of livestock management practices on contact networks. Free-ranging wild pigs (Sus scrofa) in North America are an invasive, socially-structured species with an expanding distribution that pose a threat to livestock health given their potential to transmit numerous livestock diseases, such as pseudorabies, brucellosis, trichinellosis, and echinococcosis, among many others. Our objective in this study was to quantify …

Cedarbush Creek Dredge Channel Data Report, Donna A. Milligan, C. Scott Hardaway Jr., Christine A. Wilcox, Cameron Green

Reports

Cedarbush Creek is located in Gloucester County, Virginia. It is a long, but narrow creek that empties into the York River. The mouth is a wide embayment, but farther north, the creek narrows to about 400 feet wide and extends for about 1.5 miles to its marshy headwaters. Cedarbush Creek has never been dredged, but due to shoaling within the creek, it needs dredging to accommodate vessel traffic. Oliver’s Landing, located near the mouth of Cedarbush Creek, is a working waterfront that supports commercial and recreational boaters in Gloucester. (...)

Data Collection At Fifteen Selected Creeks In Support Of Shallow Water Dredging On Virginia’S Middle Peninsula - Methods & Data Report, Nicholas J. Dinapoli, Donna A. Milligan, C. Scott Hardaway Jr., Christine A. Wilcox, Cameron Green, Scott Lerberg, Eduardo J. Miles, Alex Demeo, George Brooks

Reports

Federal funding has been historically available for the Army Corps of Engineers for shallow draft navigation projects. However, past and recent subsidies have not provided ample funding at levels to sustain maintenance dredging for the 17 federal navigation channels on the Middle Peninsula. Further, funding for maintenance of non-federal channels has been historically neglected by the Commonwealth of Virginia until the Virginia General Assembly established the Virginia Waterway Maintenance Fund in 2018. For the past decade the Middle Peninsula Chesapeake Bay Public Access Authority, the Middle Peninsula Planning District Commission and its member jurisdictions, and the Virginia Institute of Marine …

Parrotts Creek Dredge Channel Data Report, Donna A. Milligan, C. Scott Hardaway Jr., Christine A. Wilcox, Cameron W. Green

Reports

Parrotts Creek is located along the Rappahannock River in Middlesex County, VA. The mouth of the creek is about 850 ft wide, but just inside the mouth of the creek, a spit extends from the upland narrowing the creek to about 300 ft before it widens again. Overall, this is a relatively short, undeveloped creek. Most of the development (wharfs, ramps, piers) occur near the mouth. Most of the more inland areas of the creek are surrounded by woodland. The federally-authorized channel was established in 1955 due to the presence of a public ramp and landing area, as well as …

Living Shoreline Design Guidelines For Shore Protection In Virginia’S Estuarine Environment, C. Scott Hardaway Jr., Donna A. Milligan, Karen Duhring

Reports

The Chesapeake Bay has about 6.5 million people living in its coastal counties and much of the shoreline is privately-owned. For communities along the shore, the continual shore retreat may be a problem. When land along the shore show signs of erosion, property owners tend to address it.

These guidelines are meant to address the need to educate consultants, contractors, and other professionals in the use of living shoreline strategies. It provides the necessary information to determine where they are appropriate and what is involved in their design and construction. The guidelines focus on the use of created marsh fringes …

Sources And Aftermaths Of Pipeline Related Leaks And Spills, Justin Smith

Symposium of Student Scholars

The escape of oil and other hazardous materials have been shown to pollute and destroy ecosystems. As an aspiring chemist, I am adamant about the secure handling and transportation of oil and other hazardous materials. In the past, researchers have concentrated on oil’s high viscosity. Oil’s high viscosity physically smothers wildlife, affecting their ability to continue critical functions such as respiration, feeding, and thermoregulation. My research focuses on the source of these oil spills, as well as natural gas leaks, for the purpose of risk assessment. In addition, I compare recovery efforts based on the cause of the leak/spill, the …

Continental-Scale Network Reveals Cross-Jurisdictional Movements Of Sympatric Sharks With Implications For Assessment And Management, Charlie Huveneers, Yuri Niella, Michael Drew, Rory Mcauley, Paul A. Butcher, Victor Peddemors, Daniela Waltrick, Chris Dowling, Silas Mountford, Ian S. Keay, Matias Braccini

Fisheries Research Articles

Understanding the movement ecology of marine species and connectivity of populations is required for effective fisheries management. This is especially the case for species with wide-ranging distributions for which movement can span across several jurisdictions with different management regulations. We used the Australian national network of acoustic receivers facilitated by the Integrated Marine Observing System (IMOS) to describe the extent and frequency of movements for two large epipelagic shark species, the bronze whaler (Carcharhinus brachyurus) and dusky shark (Carcharhinus obscurus). A total of 210 sharks (117 bronze whalers and 93 dusky sharks) were tracked for a …

Monitoring Agroecosystem Productivity And Phenology At A National Scale: A Metric Assessment Framework, Dawn M. Browning, Eric S. Russell, Guillermo E. Ponce-Campos, Nicole Kaplan, Andrew D. Richardson, Bijan Seyednasrollah, Sheri Spiegal, Nicanor Saliendra, Joseph G. Alfieri, John Baker, Carl Bernacchi, Brandon T. Bestelmeyer, David Bosch, Elizabeth H. Boughton, Raoul K. Boughton, Pat Clark, Gerald Flerchinger, Nuria Gomez-Casanovas, Sarah Goslee, Nick M. Haddad, David Hoover, Abdullah Jaradat, Marguerite Mauritz, Gregory W. Mccarty, Gretchen R. Miller, John Sadler, Amartya Saha, Russell L. Scott, Andrew Suyker, Craig Tweedie, Jeffrey D. Wood, Xukai Zhang, Shawn D. Taylor

School of Natural Resources: Faculty Publications

Effective measurement of seasonal variations in the timing and amount of production is critical to managing spatially heterogeneous agroecosystems in a changing climate. Although numerous technologies for such measurements are available, their relationships to one another at a continental extent are unknown. Using data collected from across the Long-Term Agroecosystem Research (LTAR) network and other networks, we investigated correlations among key metrics representing primary production, phenology, and carbon fluxes in croplands, grazing lands, and crop-grazing integrated systems across the continental U.S. Metrics we examined included gross primary productivity (GPP) estimated from eddy covariance (EC) towers and modelled from the Landsat …

The Economics Of Battery Storage For Residential Solar Customers In Southern California, John B. Broughton, Prashanth U. Nyer, Candace E. Ybarra

Business Faculty Articles and Research

Battery storage coupled with solar panels became a consideration after the original net metering program in California (NEM 1.0) ended and gave way to the current net metering program (NEM 2.0). Under NEM 2.0, battery storage gives customers under time-of-use (TOU) rate plans the ability to store the excess electrical energy generated by their panels during sunlight hours (when electricity usage and resale rates are low) and then use that energy in the evening when rates are significantly higher. This reduces the amount of expensive electricity that the customer would have to purchase from the grid. It is widely expected …

Patterns And Drivers Of Introgression In Louisiana's Largemouth Bass (Micropterus Salmoides) Stocks, Colleen E. Walsh

LSU Master's Theses

In the southeastern U.S., populations of the popular sportfish Largemouth Bass (Micropterus salmoides; LMB) are often stocked with Florida Largemouth Bass (Micropterus salmoides floridanus; FLMB) to develop FLMB ancestry in systems north of its native peninsular Florida range, with the ultimate goal of creating trophy fisheries. Stock enhancement through introgression of FLMB ancestry can increase maximum potential size and growth in receiving populations, potentially through hybrid vigor of intergrade subspecies. We collected 60 fish and habitat data from six Louisiana lakes from 2018-2020 (N=360), quantified level of introgression of FLMB with fragment analysis at 12 microsatellite loci confirmed for subspecific …

Summer To Autumn Population Of Wild Eumaeus Atala On The Ft. Lauderdale Campus Of Nova Southeastern University, Alexandra M. Lens

Mako: NSU Undergraduate Student Journal

Eumaeus atala is an endangered tropical butterfly native to the Caribbean and some parts of Florida, USA. Following population reductions primarily due to habitat loss, E. atala populations are now increasing due to conservation efforts of its cycad host plants, especially Zamia integrifolia (coontie). The purpose of this study was to observe, document, and measure the population of wild E. atala on the Ft. Lauderdale, Florida campus of Nova Southeastern University where landscaping use of host plants supports a natural population of E. atala. Forty-four host plants located in two different sites were observed for 14 weeks. One site …

Effects Of Hunter Movement And Habitat Use On Observation Rate Of White-Tailed Deer (Odocoileus Virginianus), Alyssa N. Meier

Biology Theses, Dissertations, and Student Creative Activity

Hunting by humans is the primary tool for population control for many ungulate species across the United States, including white-tailed deer (Odocoileus virginianus). Previous research has focused primarily on the effects of hunting on prey behavior while neglecting the potential effects hunter behavior has on the probability of harvest success. I examined the influence of hunter movement and habitat use across the landscape on observation rate of white-tailed deer. During the 2008 and 2009 Oklahoma hunting seasons, we recorded GPS and observation data of 83 individual hunters over 487 total hunts. Hunters that moved non-linearly through forested cover at a …